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Family Rubiaceae
Hedyotis costata
Exallage cristata (Willd.) Nandikar & K.C.Kishor
Ma er cao

Scientific names Common names
Exallage costata (Roxb.) Bremek.        No common names found
Exallage cristata (Willd.) Nandikar & K.C.Kishor         
Exallage lineata (DC.) Bremek.         
Hedyotis caerulea (Blume) Korth.                       
Hedyotis capituliflora Miq.                     
Hedyotis costata (Roxb.) Kurz                     
Hedyotis vestita R.Br. G.Don                     
Matabolos caeruleus Blume                     
Matabolos lineatus DC.                     
Oldenlandia caerulea (Blume) Kuntze                   
Oldenlandia costata (Roxb.) K.Schum.                 
Oldenlandia vestita (R.Br. ex Don) Drake               
Spermacoce costata Roxb.                     
Spermacoce cristata Willd.                     
Hedyotis costata (Roxb.) Kurz is a synonym of Exallage cristata.
Exallage cristata (Willd.) Nandikar & K.C.Kishor is an accepted species. KEW: Plants of the World Online

Other vernacular names
CHINESE: Ma er cao.

Gen info
- Exallage is a genus of flowering plants in the family Rubiaceae. The genus is found from tropical and subtropical Asia to the southwestern Pacific.
- Hedyotis costata is a genus of flowering plants in the family Rubiaceae. It comprises about 115 species.
- Hedyotis, named by Linnaeus in 1753, derives from two Greek words: hedys meaning "sweet
" and otos meaning "ear", referring to the sweet-scented ear-shaped leaves of some species. (3)
- An understudied plant.
- The Hedyotis-Oldenlandia complex is one of the most enigmatic groups in Rubiaceae due to a wide range of morphological diversity observed and long-standing taxonomic complexity with conflicting generic delimitations. (5)

Botany
Hedyotis costata is a rather coarse, spreading, diffuse, villous herb 30 to 60 centimeters long. Leaves are petioled, elliptic-lanceolate, 5 to 11 centimeters long, and 1 to 3 centimeters wide. Cymes are villous, small, borne in the axils of the leaves, and furnished with short stalks under 1.5 centimeters long. Flowers are very small, white, and crowded in small heads. Fruit is a capsule, small, rounded and hairy.

• Herbs, annual or perennial, diffusely branched, weak to clambering or prostrate, to 60 cm tall; stems flattened, 4-angled, or subterete, sometimes sulcate, densely villous or hirsute with trichomes drying golden yellow. Leaves subsessile to petiolate; petiole 1-10 mm, densely hirtellous to villous; blade drying membranous to papery, lanceolate, elliptic-lanceolate, or elliptic, 2-8 × 0.5-3 cm, adaxially sparsely to densely strigose and/or strigillose to hirsute on lamina and moderately to densely pilosulous on principal veins, abaxially sparsely strigillose to glabrescent on lamina and densely strigose to hirsute on principal veins, base cuneate to obtuse and frequently decurrent, apex acute to usually acuminate; secondary veins 4-6 pairs; stipules fused to petiole bases, truncate to triangular, 2-4 mm, moderately to densely hirtellous, hirsute, or villous, with 3-5 linear or setose lobes 2-5 mm. Inflorescences axillary, congested-cymose, capitate, or fasciculate, 5-20 mm, densely hirtellous to hirsute, several to many flowered, pedunculate; peduncles 1-3 per leaf axil, 2-12 mm; bracts subulate, to 1 mm. Flowers sessile or subsessile, distylous. Calyx densely hirtellous to villosulous; hypanthium portion turbinate to obconic, ca. 0.5 mm; limb lobed nearly to base; lobes lanceolate to triangular, 0.3-1 mm. Corolla white or purple, tubular to funnelform, outside puberulent to pilosulous; tube 1.2-1.5 mm, pubescent in throat; lobes narrowly elliptic to triangular, 0.8-1.2 mm. Anthers 0.5-1 mm, exserted. Stigma 0.3-0.6 mm, exserted. Fruit indehiscent, subglobose, 0.8-1.5 mm in diam., pilosulous; seeds 6-8, black, trigonous. (Flora of China)

Distribution
- Native to the Philippines. (2) (4)
- In open grasslands and thickets at low altitudes in Rizal, Laguna, Camarines, and Sorsogon Provinces in Luzon; and in Culion, Busuanga, Palawan, and Mindanao.
- Also
native to Assam, Bangladesh, Cambodia, East Himalaya, Jawa, Laos, Malaya, Myanmar, Nepal, New Guinea, Nicobar Is., Thailand, Vietnam. (2)

Parts used
Roots.

Uses

Folkloric
- Roots boiled and made into a lotion for rheumatism.
- In Chinese medicine, used for malaria, hepatitis, rheumatism, conjunctivitis. The plant is an ingredient in a polyherbal formulation for atrophic rhinitis. (1)

Studies
Herbal Mixture for Atrophic Rhinitis:
I
nvention relates to a traditional Chinese medicine for the treatment of atrophic rhinitis prepared from a mixture of medicinal materials viz., folium eucalypti, sophora fruit, wrinkled giant hyssop herb, saffron crocus, fortune firethorn, athelenota ananas and Hedyotis costata. (1)

Availability
Wild-crafted.

Update June 2025 / June 2020 / March 2017
Updated November 2015


                                                  PHOTOS / ILLUSTRATIONS
IMAGE SOURCE: Hedyotis costata / Subject Database of China Plant /http://www.naturemuseum.net/album/ShowPhoto.aspx?photoid=2671c974-77ca-44f9-b304-0f16660a06f7> / click on image to go to source pages / Helixcn
OTHER IMAGE SOURCE: Hedyotis costata / © Jingxin Liu / Non-commercial use / Click on image or link to go to source pages / Virboga: The Virtual Botanic Garden
Additional Sources and Suggested Readings
(1)
Traditional Chinese medicine for treating atrophic rhinitis / CN 103006933 A
(2)
Exallage cristata / KEW: Plants of the World Online
(3)
Hedyotis / Wikipedia
(4)
Rubiaceae: Exallage cristata / Co's Digital Flora of the Philippines
(5)
The Hedyotis-Oldenlandia complex (Rubiaceae: Spermacoceae) in Asia and the Pacific: Phylogeny revisited with new generic delimitations / Suman Neupane, Steven Dessein, Timothy J Motley et al / TAXON, 2015; 64(2): pp 299-322 / DOI: 10.12705/642.8
(6)
Exallage / Wikipedia

DOI: It is not uncommon for links on studies/sources to change. Copying and pasting the information on the search window or using the DOI (if available) will often redirect to the new link page. (Citing and Using a (DOI) Digital Object Identifier)

                                                            List of Understudied Philippine Medicinal Plants
                                          New plant names needed
The compilation now numbers over 1,500 medicinal plants. While I believe there are hundreds more that can be added to the collection, they are becoming more difficult to find. If you have a plant to suggest for inclusion, native or introduced, please email the info: scientific name (most helpful), local plant name (if known), any known folkloric medicinal use, and, if possible, a photo. Your help will be greatly appreciated.

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